It's sometimes hard to visualize how things are on others' boats - I'm wondering if the main deck drain you're referencing is the scupper, or some other FSH drain that's not on the runabout's. As for the floor locker, that sounds like the FSH equivalent of the ski locker. If that has the white 3-rib vinyl seal all around it, save your efforts on trying to tighten the hatch to it, it's a POS that has very little adhesion, in addition to very little sealing area, from what I've seen.
I had another issue with the drain plug assembly inside the ski locker that I fixed as well, that required replacing the whole fitting. There was no sealant underneath the fitting whatsoever, and they boogered up the fitting when they screwed it in. I had to cut the fitting just to be able to unscrew it for removal. I'm hoping it proves to be water-tight once I can get my boat on the water.
View attachment 150183
This isn't the the exact fitting, but this is the manufacturer (Amarine), and the type of fitting - fully threaded thru-hull fitting. They no longer show the 3/8" fitting at this link, and this is where my Amazon order points to, so I know they USED to have it available.
Amarine-made Straight Thru-Hull Valve Fitting Fully Threaded Stainless Steel 316 Boat Thru Hull for 1-1/2 Inch Interior Diameter Hose Thru Drain Boat Hose - - Amazon.com
www.amazon.com
Check the first post out again - I edited it to post a link to another 3/8" fitting, if that's what yours requires. I purchased a 1/2" fitting from that vendor for my cleanout tray drain, they make a quality product as well, I wouldn't anticipate any issues with the smaller fitting.